none Cochylis caulocatax Razowski was previously known only from the type series, 2 males, from Venezuela. The adults, male and female genitalia, larva, and pupa are described and illustrated. Larvae feed on stamens of Eustoma grandiflorum (Raf.) Shinners (Gentianales: Gentianaceae) in early instars, then burrow into ovaries feeding on developing seeds and supporting tissues. The moth is known from north central Kentucky (USA), southeastern Texas, eastern Mississippi, south central Florida, and central Venezuela. Two generations per year occur in the Nearctic: 1) mid-June through July and 2) September to mid-October. Collections from Venezuela were in mid-February. Two parasites were reared, a braconid, Bracon sp., and a chalcidid, Spilochalcis sanguiniventris (Cresson).